We all know the Ur-Quan were easily controlled by the Dnyarri, but how did the Dnyarri cause the Ur-Quan to separate into two different sub-species? Was it genetic modification, breeding, or something completely different?
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By this point, the Dnyarri had used genetic manipulation to split the Ur-Quan into two sub-species the Green Ur-Quan -- scientists, technicians, and administrators who were responsible for maintaining the limited infrastructure of the Dnyarri civilization and the Black Ur-Quan, who filled the ranks of basic laborer and combat soldier.
I think it would be easier to determine this by seeing what the original race looked like before the split was created.
My guess is it was a matter of recoding the DNA is some shape or form (figure similar technology along the lines of what the Umgah use.) Most likely through a combination of chemical and telepathic manipulation followed by breeding the old race out with the new ones.
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Of course there are fates worse then death......after all.....you could always make me angry